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to run to seed at a great distance from my true Sprouts, which the extent of my garden allowed me easily to do. The second sowing brought them back a good deal to their true character; the plants yielded small cabbages regularly at each axil, but not generally full or compact, and they did not shoot a second time, as the true sort does. I again suffered these to run to seed, using the same precaution of keeping them by themselves. I sowed the seed, and this time the plants were found to have entirely recovered their original habits, their head, and rich produce." (Hort. Trans., iii. 197.) I must confess, however, that, although the passage merits quotation, for the sake of exciting attention to the subject, it appears to me very doubtful whether the case has been fully, if correctly, stated.

CHAP. XVIII.

OF THE IMPROVEMENT OF RACES.

WHAT has been stated in the preceding chapter, concerning the preservation of the races of domesticated plants, is in some measure applicable to their improvement; because the very means employed to preserve those peculiarities of habit,

which render them valuable, will, from time to time, be the cause of still more valuable qualities making their appearance. There are, however, other points of great importance on which the gardener has dependence.

A fixed improvement in the quality of the produce of a plant can only be obtained in one of two ways; either directly, by accidental variations in itself, or indirectly, by the process of muling.

Direct alterations in the quality of seedling plants often occur from no apparent cause, just as those accidental changes, called "sports," in the colour or form of the leaves, flowers, or fruit, of one single branch of a tree, occasionally break out, we know not why. Of these things, physiology can give no account; but it is known that, when such sports appear, they indicate a permanent constitutional change in the action of the limb thus affected, which changes may be sometimes perpetuated by seed, and always by propagation of the limb itself, when propagation is practicable. It is in this way that many of our fruits have probably, and several of the Chinese Chrysanthemums have certainly, been obtained. It was apparently thus that the Nectarine emanated from the Peach. It is possible that many new forms of shrubs might be procured by keeping these facts in view, and that climbers might be deprived of their climbing habits; for it is known that the handsome evergreen bush called

the Tree Ivy, which grows erect, with scarcely the least tendency to climb, has been procured by propagating the fruit-bearing branches of trees of considerable age.

But we are by no means destitute of the power of procuring, with considerable certainty, improved varieties, by an application to practice of physiological principles. In the last chapter has been shown the importance of securing the production of seed by plants in the most healthy state possible, because a robust parent is likely to afford a progeny of similar habits to itself. In annuals, however, this is apparently restrained within narrower limits than in woody plants, from the great difficulty of fixing a new peculiarity in the former, and the facility with which it may be effected in the latter case, by means of buds, cuttings, grafts, and similar modes of propagation. The great object of the scientific gardener who desires to improve the varieties of plants upon principle will be, then, by artificial means, to bring the parent from which seed is to be saved as near as possible to that state at which he desires the seedling to arrive.

It is well known that the abstraction of fruit and flowers augments the vigour of the branches, or of the parts connected with them, and that the removal from the former of any part which takes up a portion of the food employed in the support of the flowers increases their efficiency. Thus those

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varieties of the Potato, which will neither flower nor fruit otherwise, may be made to do both by stopping the developement of tubers; and, on the other hand, the size and weight of the tubers themselves are increased by preventing the formation of flowers and fruit. The course, then, to take, in obtaining the largest possible tubers in a new variety of the Potato, would be, in the first place, to effect that end temporarily, but during several successive seasons, by abstracting all the flowers and fruit, and by such other means as may suggest themselves; and then to obtain the most perfect seed possible by a destruction of the tubers during the season when seed is finally to be saved. Mr. Knight found, in raising new varieties of the Peach, that, when one stone contained two seeds, the plants these afforded were inferior to others. The largest seeds, obtained from the finest fruit, and from that which ripens most perfectly and most early, should always be selected (Hort. Trans., i. 39.); and, in his incessant efforts to obtain new varieties of fruit of other genera, he had reason to conclude that the trees, from blossoms and seeds of which it is proposed to propagate, should have grown at least two years in mould of the best quality; that during that period they should not be allowed to exhaust themselves by bearing any considerable crop of fruit; and that the wood of the preceding year should be thoroughly ripened

(by artificial heat when necessary) at an early period in the autumn; and, if early maturity in the fruit of the new seedling plant is required, that the fruit, within which the seed grows, should be made to acquire maturity within as short a period as is consistent with its attaining its full size and perfect flavour. Those qualities ought also to be sought in the parent fruits, which are desired in the offspring; and he found that the most perfect and vigorous progeny was obtained, of plants as of animals, when the male and female parent were not closely related to each other. (See the Horticultural Transactions, i. 165.)

There are no processes known to the cultivator so efficacious in producing new varieties as that adverted to in the last paragraph, that is to say, muling or cross breeding (88.); and it is to these operations, more than to anything else, that we owe the beauty and excellence of most of our garden productions; more, however, I think, to cross breeding than to muling. It was entirely by the first of these processes that have been so greatly multiplied and improved our fruits for the dessert, and the gay flowers that adorn our gardens. The Pelargonium, the Calceolaria, the Dahlia, the Verbena, and a thousand others - what would they be, but simple wild flowers, without the power of man exercised in this way? "To the cultivators of ornamental

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